Running through streams does let a little water in, but the upper and feet dry quickly. The upper has average breathability up front. There are two layers to the upper, with a lighter mesh on top and a perforated but thicker mesh underneath. In the midfoot and forefoot, there is more open mesh with overlays surrounding the foot. The upper is thicker in the rear part of the shoe and is reinforced both with mesh and a plastic overlay. This immediately disappears while running and I have never once noticed it even on technical terrain. There is a low external heel counter/extended side wall around the heel that I feel some pressure into my rear heel bone while standing and walking when I put the shoes on. The heel collar also fits a bit snug, which seems to narrow at the top. There is a flexible heel counter that wraps around both the medial and lateral side of the heel. The heel fits fairly normal to a hair snug. When it opens, both parts open to provide a bit more volume. The fit is fairly normal to slightly snug from the midfoot to forefoot of the shoe when the FlipRelease is closed. The On Cloudultra fits true to size in my normal US men's size 10.
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